'''Alectrosaurus''' (
IPA: ; "unmarried lizard") is a
genus of
tyrannosauroid theropod dinosaur from the
Late Cretaceous Period of
Inner Mongolia. It was a
bipedal carnivore with a body shape similar to its much larger relative, ''
Tyrannosaurus rex''. ''Alectrosaurus'' was much smaller though, most likely less than 17 feet (5 meters) long.
The generic name ''Alectrosaurus'' can also be translated as "alone lizard," and is derived from the (
Greek words ''alektros'' ("unmarried") and ''sauros'' ("lizard"). At the time of its discovery, it was unlike any other Asian carnivore known. There is one named
species (''A. olseni''), which is named in honor of George Olsen, who discovered the first specimens in
1923 on the third
American Museum of Natural History expedition to Mongolia. Both genus and species were named by
American paleontologist Charles Gilmore in
1933.
History of Discovery
The
holotype, or original specimen, of ''Alectrosaurus'' was a hindlimb discovered in the
Iren Dabasu Formation of the
Inner Mongolia Autonomous Region (Nei Mongol Zizhiqu) of the
People's Republic of China.
[Gilmore, C.W. (1933). On the dinosaurian fauna of the Iren Dabasu Formation. ''Bulletin of the American Museum of Natural History'' '67':23-78.] The age of this
geologic formation is not clear, but is commonly cited as the
Campanian stage of the Late
Cretaceous Period, about 83 to 74 million years ago.
More material, including comparable hindlimb material as well as
skull and shoulder elements, has been referred to ''Alectrosaurus''. These
fossils were found in the
Bayan Shireh Formation of
Outer Mongolia, a formation which is also of uncertain age.
[Perle, A. (1977). [On the first finding of ''Alectrosaurus'' (Tyrannosauridae, Theropoda) in the Late Cretaceous of Mongolia.] ''Problemy Geologii Mongolii'' '3':104-113. [In Russian]] It may possibly extend into the early Campanian, but recent estimates suggest it was deposited from
Cenomanian through
Santonian times.
[Hicks, J.F., Brinkman, D.L., Nichols, D.J., and Watabe, M. (1999). Paleomagnetic and palynological analyses of Albian to Santonian strata at Bayn Shireh, Burkhant, and Khuren Dukh, eastern Gobi Desert, Mongolia. ''Cretaceous Research'' '20'(6): 829-850.] Iren Dabasu and Bayan Shireh dinosaur faunas are similar (but see,
[van Itterbeecka, J., Horne, D.J., Bultynck, P., and Vandenbergh, N. (2005). Stratigraphy and palaeoenvironment of the dinosaur-bearing Upper Cretaceous Iren Dabasu Formation, Inner Mongolia, People's Republic of China. ''Cretaceous Research'' '26':699-725.] which claims the Iren Dabasu is probably
Campanian-
Maastrichtian in age and possibly correlated with the
Nemegt Formation) so it is not surprising that a species of ''Alectrosaurus'' would be found there.
Furthermore, several more partial skeletons may have been found in both Inner and Outer Mongolia.
[Currie, P.J. (2001). Theropods from the Cretaceous of Mongolia. In: Benton, M.J., Shishkin, M.A., Unwin, D.M., and Kurochkin, E.N. (Eds.). ''The Age of Dinosaurs in Russia and Mongolia''. Cambridge University Press:Cambridge, 434-455. ISBN 052154582X.] These remain undescribed as of early 2007.
Taxonomy
''Alectrosaurus'' is undoubtedly a
tyrannosauroid, but due to its fragmentary nature, there is presently very little confidence in restoring its relationships with other tyrannosauroids and many recent
cladistic analyses have omitted it altogether. One study recovered ''Alectrosaurus'' at no less than eight equally
parsimonious positions in a tyrannosauroid cladogram.
[Holtz, T.R. (2004). Tyrannosauroidea. In: Weishampel, D.A., Dodson, P., and Osmólska, H. (Eds.). ''The Dinosauria'' (2nd Edition). University of California Press:Berkeley, 111-136. ISBN 0-520-24209-2.]
''Alectrosaurus'' was originally characterized as a long-armed theropod, but this was due to the mistaken association of
segnosaur forelimb elements.
[Mader, B.J., and Bradley, R.L. (1989). A redescription and revised diagnosis of the syntypes of the Mongolian tyrannosaur ''Alectrosaurus olseni''. ''Journal of Vertebrate Paleontology'' '9'(1):41-55.] The remaining material represents the hindlimb of a true tyrannosauroid, although characterized by the low ratio between the length of its tibia and femur, meaning that both bones are about the same size, as opposed to most other tyrannosauroids, where the tibia is usually longer. The hindfoot (and ankle) are also closer in size to the tibia than most tyrannosauroids, where the foot is usually longer.
The Bayan Shireh material may or may not belong to this genus, and needs further study. One cladistic analysis showed that the two sets of specimens group together exclusive of any other taxa, so they are probably at least closely related, if not the same species.[Holtz, T.R. (2001). The phylogeny and taxonomy of the Tyrannosauridae. In: Tanke, D.H., and Carpenter, K. (Eds.). ''Mesozoic Vertebrate Life''. Indiana University Press:Bloomington, 64-83. ISBN 0-253-33907-3.]
References