DICOTYLEDON

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Young castor oil plant showing its prominent two embryonic leaves (cotyledons), that differ from the adult leaves

'Dicotyledons', or '"dicots"', is a name for a group of flowering plants whose seed typically contains two embryonic leaves or cotyledons. There are around 199,350 species within this group [1]. Flowering plants that are not dicotyledons are monocotyledons, typically having one embryonic leaf.
The dicotyledons no longer are regarded as a "good" group, and the names "dicotyledons" and "dicots" are no longer to be used at least in a taxonomic sense. The vast majority of the former dicots, however, form a monophyletic group called the eudicots or tricolpates. These may be distinguished from all other flowering plants by the structure of their pollen. Other dicotyledons and monocotyledons have monosulcate pollen, or forms derived from it, whereas eudicots have tricolpate pollen, or derived forms, the pollen having three or more pores set in furrows called colpi.
Traditionally the dicots have been called the Dicotyledones (or Dicotyledoneae), at any rank. If treated as a class, as in the Cronquist system, they may be called the Magnoliopsida after the type genus ''Magnolia''. In some schemes, the eudicots are treated as a separate class, the Rosopsida (type genus ''Rosa''), or as several separate classes. The remaining dicots (palaeodicots) may be kept in a single paraphyletic class, called Magnoliopsida, or further divided.

The following lists are of the orders formerly placed in the dicots, giving their new placement in the APG-system and that under the older Cronquist system, which is still in wide use.






'APG II' 'Cronquist system'

Amborellaceae
Chloranthaceae
Nymphaeaceae [+ Cabombaceae]
Austrobaileyales
Ceratophyllales
;magnoliids

Canellales

Laurales

Magnoliales

Piperales
;eudicots

Buxaceae [+ Didymelaceae]

Sabiaceae

Trochodendraceae [+ Tetracentraceae]

Proteales

Ranunculales

★ ;core eudicots


Aextoxicaceae


Berberidopsidaceae


Dilleniaceae


Gunnerales


Caryophyllales


Santalales


Saxifragales


★ ;rosids



Aphloiaceae



Geissolomataceae



Ixerbaceae



Picramniaceae



Strasburgeriaceae



Vitaceae



Crossosomatales



Geraniales



Myrtales



★ ;eurosids I




Zygophyllaceae [+ Krameriaceae]




Huaceae




Celastrales




Cucurbitales




Fabales




Fagales




Malpighiales




Oxalidales




Rosales



★ ;eurosids II




Tapisciaceae




Brassicales




Malvales




Sapindales


★ ;asterids



Cornales



Ericales



★ ;euasterids I




Boraginaceae




Icacinaceae




Oncothecaceae




Vahliaceae




Garryales




Gentianales




Lamiales




Solanales



★ ;euasterids II




Bruniaceae




Columelliaceae [+ Desfontainiaceae]




Eremosynaceae




Escalloniaceae




Paracryphiaceae




Polyosmaceae




Sphenostemonaceae




Tribelaceae




Apiales




Aquifoliales




Asterales




Dipsacales
Note: "+ ..." = optional segregrate family, that may be split off from the preceding family.

'Magnoliopsida'
Magnoliidae (mostly basal dicots)

Magnoliales

Laurales

Piperales

Aristolochiales

Illiciales

Nymphaeales

Ranunculales

Papaverales
Hamamelidae

Trochodendrales

Hamamelidales

Daphniphyllales

Didymelales

Eucommiales

Urticales

Leitneriales

Juglandales

Myricales

Fagales

Casuarinales
Caryophyllidae

Caryophyllales

Polygonales

Plumbaginales
Dilleniidae

Dilleniales

Theales

Malvales

Lecythidales

Nepenthales

Violales

Salicales

Capparales

Batales

Ericales

Diapensiales

Ebenales

Primulales
Rosidae

Rosales

Fabales

Proteales

Podostemales

Haloragales

Myrtales

Rhizophorales

Cornales

Santalales

Rafflesiales

Celastrales

Euphorbiales

Rhamnales

Linales

Polygalales

Sapindales

Geraniales

Apiales
Asteridae

Gentianales

Solanales

Lamiales

Callitrichales

Plantaginales

Scrophulariales

Campanulales

Rubiales

Dipsacales

Calycerales

Asterales


Contents
Compared to Monocotyledons
Compared to Monocotyledons

Aside from cotyledon number, other broad differences have been noted between monocots and dicots, although these have proven to be differences primarily between monocots and eudicots. Many early-diverging dicot groups have "monocot" characteristics such as scattered vascular bundles, trimerous flowers, and non-tricolpate pollen. In addition, some monocots have "dicot" characteristics such as reticulated leaf veins.
'Seeds:' The embryo of the monocot has one cotyledon while the embryo of the dicot has two.
'Flowers:' The flower parts in monocots are multiples of three while in dicots are multiples of four or five.
'Stems:' In monocots, the stem vascular bundles are scattered, while in dicots they are in a .
'Secondary growth:' In monocots, stems rarely show secondary growth; in dicots, stems frequently have secondary growth.
'Pollen:' In monocots, pollen has one furrow or pore while in dicots they have three.
'Roots:' The roots are adventitious in monocots, while in dicots they develop from the radicle.
'Leaves:' In monocots, the major leaf veins are parallel, while in dicots they are reticulated.

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