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GENUS


The hierarchy of scientific classification

'Genus' (plural: genera) is the first part of the name of an organism used in binomial nomenclature. An example is ''Homo sapiens'', the name for the human species (Latin for "wise man") which belongs to the genus ''Homo''.
Genera are sometimes divided into subgenera (singular: subgenus).
Each genus must have a designated type species (see Type (zoology)). The generic name is permanently associated with the type specimen of its type species. Should this specimen turn out to be assignable to another genus, the genus name linked to it becomes a junior synonym, and the remaining taxa in the now-invalid genus need to be reassessed. See scientific classification and nomenclature Codes for more details of this system. Also see type genus.
The boundaries between genera are historically subjective, but with the advent of phylogenetics, it is increasingly common for all taxonomic ranks (at least) below the class level to be restricted to demonstrably monophyletic groupings, as has been the aim since the advent of evolutionary theory. Indeed, in the better-researched groups like birds and mammals, most genera are clades already.
Rules-of-thumb for delimiting a genus are outlined e.g. in Gill ''et al.'' (2005). According to these, a genus should fulfill 3 criteria to be descriptively useful:

★ monophyly - all descendants of an ancestral taxon are grouped together;

★ reasonable compactness - a genus should not be expanded needlessly; and

★ distinctness - in regards of evolutionarily relevant criteria, i.e. ecology, morphology, or biogeography; note that DNA sequences are a ''consequence'' rather than a ''condition'' of diverging evolutionarily lineages except in cases where they directly inhibit gene flow (e.g. postzygotic barriers).
Neither the ICZN nor the ICBN require such criteria for extablishment of a genus; they rather cover the formalities of what makes a description valid. Therefore, there has been for long a vigorous debate about what criteria to consider relevant for generic distinctness. At present, most of the classifications based on phenetics - overall similarity - are being gradually replaced by new ones based on cladistics (e.g., use of Reptilia and Amphibia in taxonomy is discouraged), though phenetics was only of major relevance for a comparatively short time around the 1960s before it turned out to be unworkable.
The three criteria given above are almost always fulfillable for a given clade. An example where at least one is crassly violated no matter what the generic arrangement is are the dabbling ducks of the genus ''Anas'', which are paraphyletic in regard to the extremely distinct moa-nalos. Considering them distinct genera (as is usually done) violates criterion 1, including them in ''Anas'' violates criterion 2 and 3, and splitting up ''Anas'' so that the mallard and the American black duck are in distinct genera violates criterion 3.
A genus in one kingdom is allowed to bear a name that is in use as a genus name or other taxon name in another kingdom. Although this is discouraged by both the International Code of Zoological Nomenclature and the International Code of Botanical Nomenclature there are some five thousand such names that are in use in more than one kingdom. For instance, ''Anura'' is the name of the order of frogs but also is the name of a genus of plants (although not current: it is a synonym); and ''Aotus'' is the genus of golden peas and night monkeys; ''Oenanthe'' is the genus of wheatears and water dropworts, and ''Prunella'' is the genus of accentors and self-heal.
Obviously, within the same kingdom one generic name can apply to only one genus. This explains why the platypus genus is named ''Ornithorhynchus'' — George Shaw named it ''Platypus'' in 1799, but the name ''Platypus'' had already been given to the pinhole borer beetle by Johann Friedrich Wilhelm Herbst in 1793. Names with the same form but applying to different taxa are called homonyms. Since beetles and platypuses are both members of the kingdom Animalia, the name ''Platypus'' could not be used for both. Johann Friedrich Blumenbach published the replacement name ''Ornithorhynchus'' in 1800.
A couple of the lesser-known forms of Genus are: Caltha and Carica

Contents
See also
References
External links

See also



Cladistics

Phylogenetics

Systematics

Taxonomy

Linnaean taxonomy

References


External links



Nomenclator Zoologicus: Index of all genus and subgenus names in zoological nomenclature from 1758 to 2004.

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