
Various species of reef fish in the Northwestern Hawaiian Islands.
'Marine biology' is the scientific study of
algae,
animals and other organisms that live in the
ocean or other marine or
brackish body of water. Given that in
biology many
phyla, families and genera have some species that live in the
sea and others that live on land, marine biology classifies species based on the
environment rather than on taxonomy.
Marine life represents a vast resource, providing
food,
medicine, and raw materials, in addition to helping to support
recreation and
tourism all over the world. At a fundamental level, marine life helps determine the very nature of our planet. Marine organisms produce 90% of the
oxygen we breathe and probably help regulate the earth's
climate.
Shorelines are in part shaped and protected by marine life, and some marine organisms even help create new land.
Overview

Underwater life differs greatly from land life (shown:
cuttlefish, which have no close terrestrial relatives).
Marine biology covers a great deal, from the microscopic, including
plankton and
phytoplankton, which can be as small as 0.02
micrometers and are both hugely important as the
primary producers of the sea, to the huge
cetaceans (whales) which reach up to a reported 33
meters (109
feet) in length.
The habitats studied by marine biology include everything from the tiny layers of surface water in which organisms and abiotic items may be trapped in
surface tension between the ocean and atmosphere, to the depths of the
abyssal trenches, sometimes 10,000 meters or more beneath the surface of the ocean. It studies habitats such as
coral reefs,
kelp forests,
tidepools, muddy, sandy, and rocky bottoms, and the open ocean (pelagic) zone, where solid objects are rare and the surface of the water is the only visible boundary.
A large amount of all
life on Earth exists in the oceans. Exactly how large the proportion is is still unknown. While the oceans comprise about 71% of the Earth's surface, due to their depth they encompass about 300 times the habitable volume of the terrestrial habitats on Earth.
Many species are economically important to humans, including the
food fishes. It is also becoming understood that the well-being of marine organisms and other organisms are linked in very fundamental ways. Human understanding is growing of the relationship between life in the sea and important cycles such as that of matter (such as the
carbon cycle) and of air (such as
Earth's respiration, and movement of energy through ecosystems). Large areas beneath the ocean surface still remain effectively unexplored. Scientists know more about the moon than they know about the ocean and the life in it.
Subfields
The
marine ecosystem is large, and thus there are many subfields of marine biology. Most involve studying specializations of particular species (i.e.,
phycology,
invertebrate zoology,
ichthyology).
Other subfields study the physical effects of continual immersion in
sea water and the ocean in general, adaptation to a salty environment, and the effects of changing various oceanic properties on marine life. A subfield of marine biology studies the relationships between oceans and ocean life, and global weather and environmental issues (such as
carbon dioxide displacement).
Recent marine
biotechnology has focused largely on
marine biomolecules, especially
proteins, that may have uses in medicine or engineering. Marine environments are the home to many exotic biological materials that may inspire
biomimetic materials.
An interesting branch of marine biology is
aquaculture; which some countries do a lot of in the oceans, especially
Japan.
Related fields
Marine biology is a branch of
oceanography and is closely linked to
biology. It also encompasses many ideas from
ecology.
Fisheries science and
marine conservation can be considered partial offshoots of marine biology.
Lifeforms
Microscopic life
Microscopic life undersea is incredibly diverse and still poorly understood. For example, the role of
viruses in marine ecosystems is barely being explored even in the beginning of the
21st century.
The role of
phytoplankton is better understood due to their critical position as the most numerous
primary producers on Earth. Phytoplankton are categorized into
cyanobacteria (also called blue-green algae/bacteria), various types of
algae (red, green, brown, and yellow-green),
diatoms,
dinoflagellates,
euglenoids,
coccolithophorids,
cryptomonads,
chrysophytes,
chlorophytes,
prasinophytes, and
silicoflagellates.
Zooplankton tend to be somewhat larger, and not all are microscopic. Many
Protozoa are zooplankton, including dinoflagellates,
zooflagellates,
foraminiferans, and
radiolarians. Some of these (such as dinoflaggelates) are also phytoplankton; the
plant/animal distinction often breaks down in very small organisms. Other zooplankton include
cnidarians,
ctenophores,
chaetognaths,
molluscs,
arthropods,
urochordates, and
annelids such as
polychaetes. Many larger animals begin their life as zooplankton before they become large enough to take their familiar forms. Two examples are fish
larvae and sea stars (also called
starfish).
Plants and algae
Plant life is relatively rare undersea. Most of the
niche occupied by plants on land is actually occupied by macroscopic
algae in the ocean, such as ''
Sargassum'' and
kelp which are commonly known as
seaweeds. The non algae plants that do survive in the sea are often found in shallow waters, such as the
seagrasses (examples of which are eelgrass, ''
Zostera'', and turtlegrass, ''Thalassia''). These plants have adapted to the high salinity of the ocean environment. The
intertidal zone is also a good place to find plant life in the sea, where
mangroves or
cordgrass or
beach grass might grow.
Sea kelp is very important to small sea creatures because the creatures can hide from predators.
Marine invertebrates
As on land,
invertebrates make up a huge portion of all life in the sea. Invertebrate sea life includes
Cnidaria such as
jellyfish and
sea anemone;
Ctenophora;
sea worms including
phyla Plathyhelminthes, Nemertea, Annelida, Sipuncula, Echiura, and the Phoronida;
Mollusca including
shellfish,
squid,
octopus;
Crustaceans; Porifera including
sponges,
Bryozoa,
Echinodermata including starfish; and
Urochordata -
sea squirts or
tunicates.
Fish
Fish have
evolved very different biological functions from other large organisms. Fish anatomy includes a two-chambered heart,
operculum, secretory
cells that produce
mucous,
swim bladder,
scales,
fins,
lips and
eyes. Fish breathe by extracting
oxygen from water through their gills. Fins propel and stabilize the fish in the water.
Well known fish include:
sardines,
anchovy,
tuna,
clownfish (also known as anemonefish), and
bottom fish which include
halibut and
ling cod.
Predators include
sharks and
barracuda.
Reptiles
Main articles: Marine reptile
Reptiles which inhabit or frequent the sea include
sea turtles,
marine iguanas, sea
snakes, and saltwater
crocodiles. Since all
extant marine reptiles are
oviparous and need to return to land to lay their eggs, most reptile species live on or near land, rather than in the ocean. Some
extinct marine reptiles, such as
ichthyosaurs, evolved to be
viviparous and had no requirement to return to land.
Seabirds
Seabirds are species of
Birds adapted to living in the marine environment, examples including
albatross,
penguins,
gannets, and
auks. Although they spend most of their lives in the ocean, species such as
gulls can often be found thousands of miles inland.
Marine mammals
There are five main types of marine mammals.
★
Cetaceans include
toothed whales (Suborder
Odontoceti), such as the
Sperm Whale,
dolphins, and
porpoises. Cetaceans also include
baleen whales (Suborder
Mysticeti), such as
Gray Whales,
Humpback Whales, Dall's porpoise, and
Blue Whales.
★
Sirenians include
manatees, the
Dugong, and the extinct
Steller's Sea Cow.
★
Seals (Family
Phocidae),
sea lions (Family
Otariidae - which also include the fur seals), and the
Walrus (Family
Odobenidae) are all considered
pinnipeds.
★
Sea Otters are members of the Family
Mustelidae, which includes
weasels and
badgers.
★ Finally,
Polar Bears (Family
Ursidae) are sometimes considered marine mammals because of their dependence on the sea.
Oceanic Habitats
Reefs
Reefs comprise some of the densest and most diverse habitats in the world. The best-known types of reefs are
tropical coral reefs which exist in most tropical waters; however, reefs can also exist in cold water. Reefs are built up by
corals and other
calcium-depositing animals, usually on top of a rocky outcrop on the ocean floor. Reefs can also grow on other surfaces, which has made it possible to create
artificial reefs. Coral reefs also support a huge community of life, including the corals themselves, their symbiotic
zooxanthellae, tropical fish and many other organisms.
Much attention in marine biology is focused on coral reefs and the
El Niño weather phenomenon. In
1998, coral reefs experienced a "once in a thousand years" bleaching event, in which vast expanses of reefs across the Earth died because sea surface temperatures rose well above normal. Some reefs are recovering, but scientists say that 58
% of the world's coral reefs are now endangered and predict that
global warming could exacerbate this trend.
Deep sea and trenches
The ocean is deep, very deep in some places. The deepest recorded measure to date is the
Mariana Trench, near the
Philippines, in the
Pacific Ocean at 10924 m (35838 ft). At such depths,
water pressure is extreme and there is no sunlight, but some life still exists. Small flounder (family
Soleidae) fish and shrimp were seen by the American crew of the
bathyscaphe ''
Trieste'' when it dove to the bottom in
1960.
Other notable
oceanic trenches include
Monterey Canyon, in the eastern Pacific, the
Tonga Trench in the southwest at 10,882 m (35,702 feet), the
Philippine Trench, the
Puerto Rico Trench at 8605 m (28232 ft), the
Romanche Trench at 7760 m (24450 ft),
Fram Basin in the
Arctic Ocean at 4665 m (15305 ft), the
Java Trench at 7450 m (24442 ft), and the
South Sandwich Trench at 7235 m (23737 ft).
In general, the deep sea is considered to start at the
aphotic zone, the point where sunlight loses its power of transference through the water. Many life forms that live at these depths have the ability to create their own light.
Much life centers on
seamounts that rise from the deeps, where fish and other sea life congregate to spawn and feed.
Hydrothermal vents along the
mid-ocean ridge spreading centers act as
oases, as do their opposites,
cold seeps. Such places support unique
biomes and many new
microbes and other lifeforms have been discovered at these locations.
Open ocean
The great expanse of open ocean habitat is huge, and many species can be found passing through it and living in it. The term "open ocean" usually is meant to refer to the vast stretches of water between points of land, or between undersea mounts. Contrary to popular notions the open ocean is often not the place where marine animals spend the majority of their lives. Most species simply pass through the open ocean on their ways to other places. Larger species are the main ongoing inhabitants...
Intertidal and shore
Intertidal zones, those areas close to shore, are constantly being exposed and covered by the ocean's
tides. A huge array of life lives within this zone.
Shore habitats span from the upper intertidal zones to the area where land vegetation takes prominence. It can be underwater anywhere from daily to very infrequently. Many species here are scavengers, living off of sea life that is washed up on the shore. Many land animals also make much use of the shore and intertidal habitats. A subgroup of organisms in this habitat bores and grinds exposed rock through the process of
bioerosion.
How oceanic factors affect distribution of various organisms
An active research topic in marine biology is to discover and map the
life cycles of various species and where they spend their time. Marine biologists study how the
ocean currents,
tides and many other oceanic factors affect ocean lifeforms, including their growth, distribution and well-being. This has only recently become technically feasible with advances in
GPS and newer underwater visual devices.
Most ocean life breeds in specific places, nests or not in others, spends time as juveniles in still others, and in maturity in yet others. Scientists know little about where many species spent different parts of their life cycles. For example, it is still largely unknown where
sea turtles travel. Tracking devices do not work for some life forms, and the ocean is not friendly to
technology. But these factors are being overcome in many instances.
Famous marine biologists

Jacques-Yves Cousteau, co-inventor of the
aqua-lung, is well known for popularizing marine biology.
★
Ali Abdelghany (
1944- )
Egyptian marine biologist
★ Jakob Johan
Adolf Appellöf (
1857-
1921),
Swedish marine zoologist.
★
Joseph Ayers, marine neurophysiologist and biomimetic researcher.
★
Samuel Stillman Berry (
1887-
1984), U.S. marine zoologist.
★
Henry Bryant Bigelow (
1879 –
1967), U.S. marine biologist.
★
Rachel Carson (
1907-
1964) American Marine Biologist and Author.
★
Ilham Artüz (
1924-
1993) Turkish Marine Biologist and Oceanographer
★
Carl Chun (
1852-
1914),
German marine biologist
★
Jacques-Yves Cousteau (
1910-
1997),
French marine biologist and explorer.
★
Anton Dohrn (
1840-
1909), German marine biologist.
★
Sylvia Earle (born
1935), American oceanographer.
★
Hans Hass (born
1919), Austrian marine biologist and diving pioneer
★
Gotthilf Hempel (born
1929), German marine biologist
★
Johan Hjort, Norwegian marine zoologist and one of the founders of
ICES.
★
Bruno Hofer (
1861-
1916), German fisheries scientist.
★
Hirohito (
1901-
1989), Japanese emperor and
jellyfish taxonomist
★
Uwe Kils (born
1951), German marine biologist
★
August David Krohn (
1803 –
1891), Russian/German zoologist.
★
William Elford Leach (
1790-
1836),
English zoologist and marine biologist.
★
Philip S. Lobel Professor,
Boston University Marine Program
★
Nicholai Miklukho-Maklai (
1846-
1888),
Russian marine biologist and anthropologist.
★
Sir John Murray (
1841-
1914),
Scots-
Canadian marine biologist.
★
Ed Ricketts
★
Harald Rosenthal (born
1937), German hydrobiologist known for his work in fish farming and ecology.
★
Michael Sars (1809 – 1869),
Norwegian theologian and biologist.
★
Georg Sars (
1837-
1927), Norwegian marine biologist.
★
Ruth Turner (
1915-
2000), marine biologist.
★
Robert Paine Professor, emeritus
University of Washington intertidal Ecologist
★
Charles Wyville Thompson (
1832-
1882), Scottish marine biologist
★
Charles Darwin (
1809-
1882), wrote ''Structure and Distribution of Coral Reefs'' (1842) while aboard the
HMS Beagle.
Source:
List of biologists.
See also
★
Algae
★ ''
Aliens of the Deep'' (documentary)
★
Bioerosion
★
Fish
★
Freshwater biology
★
Important publications in marine biology
★
The Marine Mammal Center
★
Ocean
★
Phycology
★
Sea
External links
★
Global Water Facts, from Environment Canada
★
Marine Biologist Career Description
★
Oceanography Image of the Day, from the Woods Hole Oceanographic Institution
★
MarineBio: an evolving online tribute to Marine Biology and all Ocean life.
★
Marine Conservation Society: UK charity for seas and wildlife.
★
SeaTurtle.org: Tracking a sea turtle.
★
Marinebiology.co.uk a marine biology resource.
★
Marine Ecology Progress Series (MEPS).
★
Marine Biological Association of the United Kingdom.
★
Marine Life Information Network for Britain and Ireland - ''MarLIN''.
★
University of Washington Libraries Digital Collections – Freshwater and Marine Image Bank An ongoing digital collection of images related to freshwater and marine topics.
★
"Creatures of the Deep," a photo gallery and Flash video of creatures from the deep ocean.
★
PISCO - Partnership for Interdisciplinary Studies of Coastal Oceans
★
The Marine Environmental Research Institute
★
The National Ocean Service
★
Get Marine Biology Jobs, Career portal dedicated exclusively to the field of Marine Biology
★
Bioerosion website at The College of Wooster
★
Wembury Marine Centre
★
White Sea Biological Station