'Molars' are the rearmost and most complicated kind of
tooth in most
mammals. In many mammals they grind food; hence the Latin name ''mola'', "
millstone".
Human molars
Adult humans have twelve molars, in four groups of three at the back of the mouth. The third (rearmost) molar in each group is called a
wisdom tooth. It is the last tooth to appear, breaking through the surface of the gum at about the age of twenty, although this varies by ethnicity.
The types of molars in the human mouth are:
★
maxillary first molars,
maxillary second molars, and
maxillary third molars
★
mandibular first molars,
mandibular second molars, and
mandibular third molars
Molars among species

Schematic representation of a tribosphenic molar. (A) Upper. (B) Lower.
Molars differ considerably from one species to another, so there are many terms describing them:
★ 'Tribosphenic': This kind is found in
insectivores and young
platypuses (adults have no teeth). Upper molars look like three-pointed mountain ranges; lowers look like two peaks and a third off to the side.
★ 'Quadrate': This kind is found in
humans and various other species. Four cusps are arranged in a rectangle; there may be a fifth.
★ 'Bunodont': The cusps, instead of being sharp peaks, are rounded hills. The entire tooth is covered in
enamel, and is most common among omnivores such as the
pig, the
bear and
humans.
★ 'Hypsodont': There is a lot of enamel and dentine above the gumline and the top of the pulp. This kind of molar is found in mammals that wear their teeth a lot, such as the
horse.
★ 'Zalambdodont': The tooth has two ridges that meet at an angle, forming the letter lambda.
★ 'Dilambdodont': Like zalambdodont, but there are two lambdas on one tooth.
★ 'Lophodont': The tooth has a few ridges perpendicular to the jaw.
★ 'Selenodont': The tooth has a crescent-shaped ridge or ridges.
★ 'Loxodont': The tooth has several parallel oblique ridges on its surface. The
elephant ''Loxodonta'' is named for this feature.
Tribosphenic molar
The
molar design that is considered one of the most important characteristics of mammals is a three-cusped shape called a 'tribosphenic molar'. This design of molar has two important features: the ''
trigonoid'', or shearing end, and the ''
talinoid'', or crushing heel. With the exception of Jurassic mammal ''
Shuotherium'', the talinoid is posterior the triginoid.
The tribosphenic design appears in all species of
mammals. In
monotremes, it seems to have developed independently, rather than from common ancestry with
marsupials and
placentals.
Additional images
External links
For pictures of various molars see
The Diversity of Cheek Teeth.