The 'pelycosaurs' (from
Greek ''pelyx'' meaning 'bowl' and ''sauros'' meaning 'lizard') were primitive Late
Paleozoic synapsid amniotes. Some species were quite large and could grow up to 3 meters or more, although most species were much smaller. They appeared during the
Late Carboniferous and reached their acme in the
early part of the Permian Period, remaining the dominant land animals for many millions of years. A few continued into the
late Permian.
At least two pelycosaur
clades independently evolved a tall sail, consisting of elongated vertebral spines: the
edaphosaurids and the
sphenacodontids. In life, this would have been covered by skin, and possibly functioned as a
thermoregulatory device or for mating display. Pelycosaur fossils have been found mainly in
Europe and
North America, although some small, late-surviving forms are known from
Russia and
South Africa.
Well-known pelycosaurs include the
genera ''
Dimetrodon'', ''
Sphenacodon'', ''
Edaphosaurus'', and ''
Ophiacodon''.
In
1940 the group was reviewed in detail and every species known at the time described (and many illustrated) in an important monograph by
Alfred Sherwood Romer and
Llewellyn Price.
Pelycosauria is a
paraphyletic taxon because it excludes the
therapsids. For that reason the term is not used in some modern books.
Eupelycosauria is used to designate the clade that includes most Pelycosaurs along with the Therapsida and the Mammals. In contrast to "Pelycosaurs", this is
monophyletic group.
Caseasauria refers to a pelycosaur side-branch or clade that did not leave any descendants.
The pelycosaurs appear to have been a group of synapsids that had direct ancestral links with the mammalia, having differentiated teeth and a developing hard palate.
Taxonomy and Phylogeny
★ 'Class
Synapsida'
★
★ 'ORDER PELYCOSAURIA'
★
★
★
★ 'Suborder
Caseasauria'
★
★
★
★ Family
Eothyrididae
★
★
★
★
★ ''
Eothyris''
★
★
★
★
★ ''
Oedaleops''
★
★
★
★ Family
Caseidae
★
★
★
★
★ ''
Angelosaurus''
★
★
★
★
★ ''
Casea''
★
★
★
★
★ ''
Caseopsis''
★
★
★
★
★ ''
Cotylorhynchus''
★
★
★
★
★ ''
Ennatosaurus''
★
★
★
★
★ ''
Knoxosaurus''
★
★
★
★
★ ''
Caseoides''
★
★
★ 'Suborder
Eupelycosauria'
★
★
★
★ Family
Varanopseidae
★
★
★
★
★ ''
Varanosaurus''
★
★
★
★
★ ''
Archaeovenator''
★
★
★
★
★ ''
Pyozia''
★
★
★
★
★ ''
Mycterosaurus''
★
★
★
★
★ ''
Mesenosaurus''
★
★
★
★
★ ''
Elliotsmithia''
★
★
★
★
★ ''
Ruthiromia''
★
★
★
★
★ ''
Aerosaurus''
★
★
★
★
★ ''
Varanodon''
★
★
★
★
★ ''
Varanops''
★
★
★
★ Family
Ophiacodontidae
★
★
★
★
★ ''
Archaeothyris''
★
★
★
★
★ ''
Baldwinonus''
★
★
★
★
★ ''
Clepsydrops''
★
★
★
★
★ ''
Limnostygis''
★
★
★
★
★ ''
Ophiacodon''
★
★
★
★
★ ''
Stereophallodon''
★
★
★
★
★ ''
Stereorhachis''
★
★
★
★ Family
Edaphosauridae
★
★
★
★
★ ''
Edaphosaurus''
★
★
★
★
★ ''
Ianthasaurus''
★
★
★
★
★ ''
Glaucosaurus''
★
★
★
★ Family
Lupeosauridae
★
★
★
★
★ ''
Lupeosaurus''
★
★
★
★
Sphenacodontia
★
★
★
★
★ ?''
Watongia''
★
★
★
★
★ ''
Haptodus''
★
★
★
★
★ ''
Palaeohatteria''
★
★
★
★
★ ''
Pantelosaurus''
★
★
★
★
★ ''
Cutleria''
★
★
★
★
★
Sphenacodontoidea
★
★
★
★
★
★ Family
Sphenacodontidae
★
★
★
★
★
★
★ ''
Ctenorhachis''
★
★
★
★
★
★
★ ''
Steppesaurus''
★
★
★
★
★
★
★ ''
Bathygnathus''
★
★
★
★
★
★
★ ''
Ctenospondylus''
★
★
★
★
★
★
★ ''
Dimetrodon''
★
★
★
★
★
★
★ ''
Neosaurus''
★
★
★
★
★
★
★ ''
Secodontosaurus''
★
★
★
★
★
★
★ ''
Sphenacodon''
★
★ 'ORDER
THERAPSIDA
★
See also
★
Mammal-like reptiles
References
★
Reisz, R. R., 1986, ''Handbuch der Paläoherpetologie – Encyclopedia of Paleoherpetology, Part 17A Pelycosauria''
Verlag Dr. Friedrich Pfeil, ISBN 3-89937-032-5
★
Romer, AS &
Price L.I (1940), Review of the Pelycosauria. ''Geol. Soc. Amer. Spec. Papers'' 28: 1-538.
External links
★
Introduction to the Pelycosaurs - at UCMP
★
Synapsida - Pelycosauria - at Palaeos