The 'strawberry' (''Fragaria'') is a genus of
plants in the family ''
Rosaceae'' and the
fruit of these plants. There are more than 20 named
species and many hybrids and
cultivars. The most common strawberries grown commercially are cultivars of the
Garden strawberry, ''Fragaria'' × ''ananassa''.
Morphology
The strawberry is an
accessory fruit; that is, the fleshy part is derived not from the ovaries which are the "seeds" (actually
achenes) but from the peg at the bottom of the
hypanthium that held the ovaries. So from a technical standpoint, the seeds are the actual fruits of the plant, and the flesh of the strawberry is modified receptacle tissue. It is whitish-green as it develops and in most species turns red when ripe.
History
The typical modern strawberry, of the genus ''Fragaria'', comes from the
Americas, and is a hybrid of both North and South American varieties. Interestingly, the crossbreeding was done in Europe to correct a mistake; the European
horticulturists had only brought female South American plants, and were forced to cross them with the North American variety in order to get fruit and seeds.
Fragaria comes from "fragans", meaning odorous, referring to the perfumed flesh of the fruit. Madam
Tallien, a great figure of the
French Revolution, who was nicknamed Our Lady of
Thermidor, used to take baths full of strawberries to keep the full radiance of her skin.
Fontenelle, centenarian writer and gourmet of the 18th century, considered his long life was due to the strawberries he used to eat.
Strawberries were considered poisonous in Argentina until the mid-nineteenth century.
Classification
There are more than 20 different ''Fragaria'' species worldwide. Key to the classification of strawberry species is recognizing that they vary in the number of
chromosomes. There are seven basic ''types'' of chromosomes that they all have in common. However, they exhibit different
polyploidy. Some species are diploid, having two sets of the seven chromosomes (14 chromosomes total). Others are tetraploid (four sets, 28 chromosomes total), hexaploid (six sets, 42 chromosomes total), octoploid (eight sets, 56 chromosomes total), or decaploid (ten sets, 70 chromosomes total).
As a rough rule (with exceptions), strawberry species with more chromosomes tend to be more robust and produce larger plants with larger berries (Darrow).
;Diploid species
★ ''
Fragaria daltoniana''
★ ''
Fragaria iinumae''
★ ''
Fragaria nilgerrensis''
★ ''
Fragaria nipponica''
★ ''
Fragaria nubicola''
★
''Fragaria vesca'' (Woodland Strawberry)
★ ''
Fragaria viridis''
★ ''
Fragaria yezoensis''
;Tetraploid species
★ ''
Fragaria moupinensis''
★ ''
Fragaria orientalis''
;Hexaploid species
★
''Fragaria moschata'' (Musk Strawberry)
;Octoploid species and hybrids
★
''Fragaria x ananassa'' (Garden Strawberry)
★
''Fragaria chiloensis'' (Beach Strawberry)
★
''Fragaria iturupensis'' (Iturup Strawberry)
★
''Fragaria virginiana'' (Virginia Strawberry)
;Decaploid species and hybrids
★
Fragaria × Potentilla hybrids
★ ''
Fragaria × vescana''
Numerous other species have been proposed. Some are now recognized as subspecies of one of the above species (see GRIN taxonomy database).
The
Mock Strawberry and
Barren Strawberry, which both bear resemblance to ''Fragaria'', are closely related species in the genus ''
Potentilla''. The
Strawberry tree is an unrelated species.
Production trends

Strawberry output in 2005
The
FAO reports that the United States was the top producer of strawberry worldwide in 2005 followed by Spain.
Pests
A number of species of
Lepidoptera feed on strawberry plants; for details see
this list.
Diseases
Etymology
The name is derived from
Old English '' which is a compound of ''streaw'' meaning "straw" and ''berige'' meaning "berry". The reason for this is unclear. It may derive from the strawlike appearance of the runners, or from an obsolete denotation of straw, meaning "chaff", referring to the scattered appearance of the achenes.
Interestingly, in other Germanic countries there is a tradition of collecting wild strawberries by threading them on straws. In those countries people find ''straw-berry'' to be an easy word to learn considering their association with straws. (In fact, in almost all other Germanic languages besides English, the name of the fruit corresponds to "Earth-Berry", so this claim is dubious.)
There is an alternative theory that the name derives from the Anglo-Saxon verb for "strew" (meaning to spread around) which was streabergen (Strea means "strew" and Bergen means "berry" or "fruit") and thence to streberie, straiberie, strauberie, straubery, strauberry, and finally, "strawberry", the word which we use today. The name might have come from the fact that the fruit and various runners appear "strewn" along the ground.
Popular etymology has it that it comes from gardeners' practice of
mulching strawberries with
straw to protect the fruits from rot (a pseudoetymology that can be found in non-linguistic sources such as the
Old Farmer's Almanac 2005). However, there is no evidence that the Anglo-Saxons ever grew strawberries, and even less that they knew of this practice.
Gallery
See also
★
Garden Strawberry
★
Epigynous berry
References and external links
★ Darrow, George M. The Strawberry: History, Breeding and Physiology. New York. Holt, Rinehart and Winston, 1966.
★
Available online.
★
List of Fragaria resources, USDA
★
GRIN Fragaria Taxonomy Database Listing of ''Fragaria'' species, also from a USDA website
★
Medicinal uses of strawberries in Armenia
★
''Fragaria chiloensis'' pictures from ''Chilebosque''
★
★
Strawberry pest management guidelines
★
Crossing of cv. 'Mieze Schindler' with cv. 'Elsanta'